1. Department of Biology, University of Nevada, Reno , Reno , NV 89557 , United States
2. Ecology, Evolution, and Conservation Biology, University of Nevada , Reno, NV 89557 , United States
3. Bureau of Land Management, Nevada State Office , Reno, NV 89502 , United States
4. Department of Botany, University of Wyoming , Laramie, WY, 82071 , United States
5. Rocky Mountain Research Station, USDA Forest Service , Moscow, ID 83843 , United States