1. State Key Laboratory of Hybrid Rice Hunan Hybrid Rice Research Center Changsha 410128 China
2. International Rice Research Institute Metro Manila 1301 Philippines
3. Guangdong Provincial Key Laboratory of High Technology for Plant Protection, Plant Protection Research Institute Guangdong Academy of Agricultural Sciences Guangzhou 510640 China
4. Department of Plant Pathology, College of Plant Protection, Nanjing Agricultural University, and Key Laboratory of Integrated Management of Crop Diseases and Pests Ministry of Education Nanjing 210095 China
5. State Key Laboratory of Ecological Pest Control for Fujian and Taiwan Crops Fujian Agriculture and Forestry University Fuzhou 350002 China
6. Department of Biochemistry and Metabolism, John Innes Centre Norwich Research Park Norwich NR4 7UH UK