1. State Key Laboratory of Tree Genetics and Breeding, School of Forestry Northeast Forestry University Harbin 150040 China
2. College of Life Sciences Northeast Forestry University Harbin 150040 China
3. College of Forest Resources and Environmental Science Michigan Technological University Houghton Michigan 49931 USA
4. Key Laboratory of Saline‐alkali Vegetation Ecology Restoration (Northeast Forestry University) Ministry of Education Harbin 150040 China