1. Department of Marine Sciences; University of Otago; PO Box 56 Dunedin 9054 New Zealand
2. Department of Botany; University of Otago; PO Box 56 Dunedin 9054 New Zealand
3. Institute for Marine and Antarctic Studies; University of Tasmania; Hobart Tasmania 7001 Australia
4. School of Biological Sciences; Monash University; Clayton Victoria 3800 Australia