1. Livestock and Wildlife Laboratory, Arid Lands Institute of Medenine University of Gabes Medenine 4119 Tunisia
2. Univ Lyon, Université Claude Bernard Lyon 1, ISARA Lyon, BioDyMIA (Bioingénierie et Dynamique Microbienne aux Interfaces Alimentaires) Research Unit, IUT Lyon 1 Bourg en Bresse 01000 France
3. Department of Food, High Institute of Applied Biology of Medenine University of Gabes Medenine 4119 Tunisia