1. School of Biological & Biomedical Sciences; Durham University; South Road Durham DH1 3LE UK
2. Department of Biological Sciences; Institute of Bioinformatics and Evolutionary Studies; University of Idaho; Moscow ID 83844-3051 USA
3. Department of Animal Science; University of California; One Shields Avenue Davis CA 95616 USA
4. Flathead Lake Biological Station; Fish and Wildlife Genomics Group; University of Montana; Polson MT 59860 USA
5. School of Aquatic and Fishery Sciences; 1122 NE Boat Street Box 355020 Seattle WA 98195-5020 USA