1. National Key Laboratory of Plant Molecular Genetics, CAS‐JIC Centre of Excellence for Plant and Microbial Science, Center for Excellence in Molecular Plant Sciences Chinese Academy of Sciences Shanghai 200032 China
2. University of Chinese Academy of Sciences Beijing 100039 China
3. Hunan University of Arts and Science Changde 415000 China
4. The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, School of Life Science Shandong University Qingdao 266237 China
5. Cell and Developmental Biology John Innes Centre, Norwich Research Park Norwich NR4 7UH UK