1. Cell and Developmental Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom
2. Computational and Systems Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom
3. Department of Molecular Microbiology, John Innes Centre, Norwich NR4 7UH, United Kingdom
4. Division of Plant Biology, The Samuel Roberts Noble Foundation, Ardmore, Oklahoma 73401