1. Key Laboratory of Plant Resource Conservation and Germplasm Innovation in Mountainous Region (Ministry of Education), College of Agricultural Sciences Guizhou University Guiyang 550025 China
2. Hubei Engineering Research Center of Viral Vector Wuhan University of Bioengineering Wuhan 430415 China
3. National Key Laboratory of Wheat and Maize Crop Science, College of Agronomy Henan Agricultural University Zhengzhou 450002 China