1. Field Science Center for Northern Biosphere Hokkaido University Sapporo Japan
2. Graduate School of Agriculture Hokkaido University Sapporo Japan
3. DKK‐Toa Yamagata Co Shinjo Japan
4. Organization for Research Promotion Iwate University Morioka Japan
5. Institute of Livestock and Grassland Science NARO Tsukuba Japan
6. Research Faculty of Agriculture Hokkaido University Sapporo Japan