1. Institute for Applied Ecology; University of Canberra; Canberra ACT 2601 Australia
2. School of Environmental and Rural Science; University of New England; Armidale New South Wales 2351 Australia
3. Finnish Environment Institute, Natural Environment Centre, Biodiversity; Oulu Finland
4. 7 Dalrymple Crescent; Pymble New South Wales 2073 Australia
5. Division of Biology; Kansas State University; Manhattan KS U.S.A.