1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of BotanyChinese Academy of SciencesKunming 650201 China
2. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations, Kunming Institute of BotanyChinese Academy of SciencesKunming 650201 China
3. College of Life SciencesUniversity of Chinese Academy of SciencesBeijing 100049 China