1. Research Centre for Ecological Change, Organismal and Evolutionary Biology Research Programme University of Helsinki Helsinki Finland
2. Finnish Environment Institute (SYKE) Helsinki Finland
3. Kainuu Centre for Economic Development, Transport and the Environment Kajaani Finland
4. Spatial Foodweb Ecology Group Department of Agricultural Sciences University of Helsinki Helsinki Finland
5. Spatial Foodweb Ecology Group Department of Ecology Swedish University of Agricultural Sciences Uppsala Sweden