1. Department of Organismic and Evolutionary Biology Harvard University Herbaria 22 Divinity Avenue Cambridge MA 02138 USA
2. Key Laboratory of Bio‐resource and Eco‐environment of Ministry of Education College of Life Sciences Sichuan University Chengdu 610064 China
3. Departamento de Ciências Biológicas Universidade Estadual de Santa Cruz Ilhéus 45.662‐900 Bahia Brazil
4. Rimba Ilmu Botanic Garden Institute of Biological Sciences University of Malaya 50603 Kuala Lumpur Malaysia
5. Department of Ecology and Evolution Stony Brook University Stony Brook NY 11794 USA
6. Department of Statistics and Institute of Bioinformatics University of Georgia Athens GA 30602 USA