1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology, Institute of Biodiversity, School of Ecology and Environmental Science, Yunnan University Kunming Yunnan China
2. School of Biology and Ecology, College of Natural Science, Forestry and Agriculture, University of Maine Orono ME USA
3. Climate Change Institute, University of Maine Orono ME USA
4. Anhui Provincial Key Laboratory of the Conservation and Exploitation of Biological Resources, College of Life Sciences, Anhui Normal University Wuhu Anhui China
5. State Key Laboratory of Plant Diversity and Specialty Crops, Wuhan Botanical Garden, Chinese Academy of Sciences (CAS) Wuhan Hubei China