1. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming China
2. CAS Key Laboratory for Plant Diversity and Biogeography of East AsiaKunming Institute of Botany, Chinese Academy of Sciences Kunming China
3. Institute of Ecology and Geobotany, School of Ecology and Environmental SciencesYunnan University Kunming China
4. Department of Plant SciencesUniversity of Oxford UK
5. The Queen's College High Street Oxford UK