1. ARC Centre of Excellence for Coral Reef Studies; School of Marine and Tropical Biology; James Cook University; Townsville Qld 4811 Australia
2. Centre for Marine Microbiology and Genetics; Australian Institute of Marine Science; PMB 3 Townsville MC Townsville Qld 4810 Australia
3. AIMS@JCU; James Cook University; Townsville Qld 4811 Australia