1. Department of Plant Pathology; China Agricultural University; Beijing 100193 China
2. State Key Laboratory for Biology of Plant Disease and Insect Pests; Institute of Plant Protection; Chinese Academy of Agricultural Sciences; Beijing 100193 China
3. Institute of Plant Virology; Key Laboratory of Plant Virology of Fujian Province; Fujian Agriculture and Forestry University; Fuzhou 350002 China
4. Department of Biological Chemistry and Crop Protection; Rothamsted Research; Harpenden AL5 2JQ UK
5. Institute of Life Science and College of Medicine; Swansea University; Swansea SA2 8PP UK