1. College of Science and Engineering; Australian Tropical Herbarium; Centre for Tropical Environmental and Sustainability Science; James Cook University; Cairns Queensland Australia
2. Department of Forest Ecology and Management; Swedish University of Agricultural Science; Umeå Sweden
3. Institute of Ecology and Earth Sciences; University of Tartum; Tartu Estonia
4. Natural History Museum and Institute of Ecology and Earth Sciences; University of Tartu; Tartu Estonia
5. College of Science and Engineering; Centre for Tropical Environmental and Sustainability Science; James Cook University; Cairns Queensland Australia