1. School of Biological Sciences; The University of Queensland; St Lucia QLD 4072 Australia
2. School of Earth and Environmental Sciences; The University of Queensland; St Lucia QLD 4072 Australia
3. The Nature Conservancy; South Brisbane QLD 4101 Australia
4. Australian Research Council Centre of Excellence for Coral Reef Studies; James Cook University; Townsville QLD 4811 Australia
5. School of Life and Environmental Sciences, Centre for Integrative Ecology; Deakin University; Burwood VIC 3125 Australia