1. Department of Ecology and Genetics, Animal Ecology Uppsala University Norbyvagen 18D, 75236 Uppsala Sweden
2. CEFE Univ Montpellier CNRS Univ Paul Valéry Montpellier 3 EPHE IRD Montpellier France
3. Department of Evolutionary Biology and Environmental Studies University of Zürich Winterthurerstrasse 190 CH‐8057 Zürich Switzerland
4. School of Biological Sciences University of East Anglia Norwich Research Park NorwichNR4 7TJUK