1. Key Lab of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education; Huazhong Agricultural University, Freshwater Aquaculture Collaborative Innovation Center of Hubei Province; Wuhan 430070 China
2. Key Lab of Freshwater Animal Breeding, College of Fisheries, Ministry of Agriculture; Huazhong Agricultural University, Freshwater Aquaculture Collaborative Innovation Center of Hubei Province; Wuhan 430070 China
3. Key Laboratory of Freshwater Fisheries and Germplasm Resources Utilization, Ministry of Agriculture; Freshwater Fisheries Research Center, Chinese Academy of Fishery Sciences; Wuxi 214081 China