1. Department of Zoology and Entomology; University of Pretoria; Pretoria 0002 South Africa
2. Forestry and Agricultural Biotechnology Institute (FABI); University of Pretoria; Pretoria 0002 South Africa
3. School of Plant Sciences; Haramaya University; Dire Dawa, PO Box 38 Ethiopia
4. Department of Biochemistry, Genetics and Microbiology, Forestry and Agricultural Biotechnology Institute (FABI); University of Pretoria; Pretoria 0002 South Africa
5. Department of Natural Resources and the Environment; University of New Hampshire; Durham NH 03824 U.S.A.