1. State Key Laboratory of Genetic Engineering and Ministry of Education Key Laboratory of Biodiversity Sciences and Ecological Engineering Institute of Plant Biology School of Life Sciences Fudan University Shanghai 200438 China
2. Shanghai Chenshan Plant Science Research Center Chinese Academy of Sciences Chenshan Botanical Garden Shanghai 201602 China
3. Key Laboratory of Plant Functional Genomics of the Ministry of Education Jiangsu Key Laboratory of Crop Genetics and Physiology/Co‐Innovation Center for Modern Production Technology of Grain Crops Yangzhou University Yangzhou 225009 China
4. Department of Biology and the Integrative Program for Biological and Genome Sciences University of North Carolina at Chapel Hill Chapel Hill NC 27599‐3280 USA
5. Lineberger Comprehensive Cancer Center University of North Carolina School of Medicine Chapel Hill NC 27599‐3280 USA
6. Center for Evolutionary Biology Institutes of Biomedical Sciences School of Life Sciences Fudan University Shanghai 200433 China