1. Key Laboratory of Plant Resources and Beijing Botanical Garden Institute of Botany, Chinese Academy of Sciences Xiangshan Beijing 100093 China
2. Beijing Key Laboratory of Grape Science and Enology Institute of Botany, Chinese Academy of Sciences Xiangshan Beijing 100093 China
3. State Key Laboratory of Systematic and Evolutionary Botany Institute of Botany, Chinese Academy of Sciences Beijing 100093 China
4. University of Chinese Academy of Sciences Beijing 100049 China
5. Shandong University of Traditional Chinese Medicine Jinan 250300 China
6. College of Forestry Inner Mongolian Agricultural University Hohhot 010019 China
7. Institute of Gardening, Hohhot Inner Mongolia Hohhot 010030 China