1. State Key Laboratory of Marine Environmental Science, Institute of Marine Microbes and Ecospheres; College of Ocean and Earth Sciences, Xiamen University; Xiamen 361102 China
2. Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystem; College of the Environment and Ecology, Xiamen University; Xiamen 361102 China
3. Department of Life Sciences; Natural History Museum; London SW7 5BD UK