1. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography Chinese Academy of Sciences Urumqi China
2. Key Laboratory of Environmental and Applied Microbiology Chinese Academy of Sciences, and Environmental Microbiology Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences Chengdu China
3. College of Resources and Environment Fujian Agriculture and Forest University Fuzhou China
4. University of the Chinese Academy of Sciences Beijing China
5. Department of Plant and Wildlife Sciences Brigham Young University Provo Utah USA