1. State Key Laboratory of Soil and Sustainable Agriculture; Institute of Soil Science, Chinese Academy of Sciences; East Beijing Road 71 Nanjing 210008 China
2. Novogene Bioinformatics Institute; Beijing 100083 China
3. Department of Biological Sciences; Seoul National University; Gwanak Seoul 151 South Korea
4. Key Laboratory of Ecosystem Network Observation and Modeling; Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences; Beijing 100101 China