1. State Key Laboratory of Microbial Technology; Shandong University; Jinan 250100 People's Republic of China
2. State Key Laboratory of Microbial Metabolism, Joint International Research Laboratory of Metabolic & Developmental Sciences, and School of Life Sciences & Biotechnology; Shanghai Jiao Tong University; Shanghai 200240 People's Republic of China
3. Shenzhen Research Institute of Shandong University; Shenzhen 518057 People's Republic of China