1. GBF Laboratory Université de Toulouse INRA Castanet‐Tolosan 31320 France
2. Key Laboratory of Bio‐Resource and Eco‐Environment of Ministry of Education College of Life Sciences Sichuan University Chengdu 610065 Sichuan China
3. Laboratory of Plant Physiology Institute of Biosciences Department of Botany Universidade de São Paulo São Paulo 11461 Brazil
4. Laboratoire de Reproduction et Développement des Plantes CNRS, INRA, ENS de Lyon, UCBL Université de Lyon Lyon 69364 France