1. Biotechnology Research Institute CAAS/Key Laboratory of Agricultural Genomics (Beijing), Ministry of Agriculture 100081 Beijing China
2. National Nanfan Research Institute (Sanya) 572022 Sanya Hainan China
3. National Centre for Plant Gene Research (Beijing), Institute of Genetics and Developmental Biology Chinese Academy of Sciences 100101 Beijing China
4. College of Advanced Agricultural Sciences University of Chinese Academy of Sciences 100039 Beijing China