1. State Key Laboratory of Crop Genetics & Germplasm Enhancement, Academy for Advanced Interdisciplinary Studies, Jiangsu Collaborative Innovation Center for Modern Crop Production Co‐sponsored by Province and Ministry Nanjing Agricultural University Nanjing 210095 China
2. National Center for Gene Research, CAS Center for Excellence in Molecular Plant Sciences Chinese Academy of Sciences Shanghai 200233 China
3. Cambridge Crop Research National Institute of Agricultural Botany (NIAB) Cambridge CB3 0LE UK
4. Earlham Institute Norwich Research Park Norwich NR4 7UH UK