1. Department of Environmental Biology; University of Rome ‘La Sapienza’; Piazzale Aldo Moro 5 00185 Rome Italy
2. Laboratoire d'Ecologie Alpine; Unité Mixte de Recherche 5533 CNRS-Université Joseph Fourier; Grenoble France
3. Department of Sciences; University ‘Roma Tre’; Viale Marconi 446 00146 Roma Italy