1. State Key Laboratory of Plant Physiology and Biochemistry, College of Life Sciences Zhejiang University Hangzhou 310027 China
2. College of Environmental and Resource Sciences Zhejiang University Hangzhou 310027 China
3. State Key Laboratory of Crop Stress Adaptation and Improvement, School of Life Sciences Henan University Kaifeng 475004 China
4. Department of Biology University of York Heslington York YO10 5DD UK