1. U.S. Fish and Wildlife Service Fort Collins Colorado USA
2. Wildlife Biology Program University of Montana Missoula Montana USA
3. Department of Biology Appalachian State University Boone North Carolina USA
4. U. S. Geological Survey, Montana Cooperative Wildlife Research Unit University of Montana Missoula Montana USA
5. Wildlife Biology Program and Office of the Vice President for Research and Creative Scholarship University of Montana Missoula Montana USA