1. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design College of Biological Sciences and Technology Beijing Forestry University No. 35, Qinghua East Road Beijing 100083 China
2. National Engineering Laboratory for Tree Breeding College of Biological Sciences and Technology Beijing Forestry University 35 Qinghua East Road Beijing China
3. Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants Ministry of Education College of Biological Sciences and Technology Beijing Forestry University 35 Qinghua East Road Beijing China
4. Department of Forest and Conservation Sciences Faculty of Forestry Forest Sciences Centre University of British Columbia Vancouver BC V6T 1Z4 Canada