1. School of Life Sciences Nanjing University Jiangsu 210093 China
2. Marine Biological Laboratory The Ecosystems Center Woods Hole MA 02543 USA
3. Department of Earth, Environmental and Planetary Sciences Brown University Providence RI 02912 USA
4. Department of Environmental Sciences University of Virginia Charlottesville VA 22903 USA
5. International Institute for Earth System Science Nanjing University Jiangsu 210093 China
6. Department of Earth and Planetary Sciences School of Engineering and Applied Sciences Harvard University Cambridge MA USA