1. State Key Laboratory of Soil and Sustainable Agriculture Institute of Soil Science Chinese Academy of Sciences East Beijing Road 71 Nanjing 210008 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Department of Biological Sciences Seoul National University Gwanak Seoul 151 Korea
4. Department of Ecology College of Urban and Environmental Sciences and Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University 5 Yiheyuan Road Beijing 100871 China
5. Key Laboratory of Adaptation and Evolution of Plateau Biota Northwest Institute of Plateau Biology Chinese Academy of Sciences 23 Xinning Road Xining 810008 China
6. Natural History Museum University of Tartu 14a Ravila Tartu 50411 Estonia