1. Department of Zoology; University of British Columbia; 6270 University Blvd Vancouver BC Canada V6T1Z4
2. Department of Botany; University of British Columbia; 6270 University Blvd Vancouver BC Canada V6T1Z4
3. Ecology and Evolutionary Biology; University of Colorado at Boulder; Ramaley N122 Boulder CO 80309-0334 USA
4. Department of Forest and Conservation Sciences; University of British Columbia; 2424 Main Mall Vancouver BC Canada V6T1Z4
5. Molecular Ecology School of Environmental and Rural Science; University of New England Armidale; Armidale NSW 2351 Australia
6. Centre for Genomic Regulation and UPF; Dr Aiguader 88 Barcelona 08003 Spain