1. School of Marine and Tropical Biology and ARC Centre of Excellence for Coral Reef Studies; James Cook University; Townsville Qld 4811 Australia
2. College of Forestry, Agriculture and Natural Resource Management; University of Hawaii at Hilo; Hilo HI 96720 U.S.A.
3. Marine Spatial Ecology Lab, ARC Centre of Excellence for Coral Reef Studies and School of Biological Sciences; University of Queensland; Brisbane Qld 4072 Australia
4. Department of Environment and Agriculture; Curtin University; Perth WA 6845 Australia