1. Department of Biological Sciences Simon Fraser University Burnaby V5A 1S6 Canada
2. Tennenbaum Marine Observatories Network Smithsonian Institution Edgewater MD 21037 U.S.A.
3. Function, Evolution and Anatomy Research (FEAR) Lab and Palaeoscience Research Centre, School of Environmental and Rural Science University of New England Armidale 2351 Australia
4. College of Science and Engineering James Cook University Townsville 4811 Australia
5. ARC Centre of Excellence for Coral Reef Studies James Cook University Townsville 4811 Australia
6. School of Aquatic and Fishery Sciences, and the Burke Museum of Natural History and Culture University of Washington Seattle WA 98105 U.S.A.