1. Centre for Ecology and Conservation, College of Life and Environmental Sciences; University of Exeter; Penryn Campus Penryn Cornwall TR10 9FE United Kingdom
2. Department of Bioscience, Terrestrial Ecology; Aarhus University; Vejlsøvej 25 8600 Silkeborg Denmark
3. School of Science and Health; Western Sydney University; Locked Bag 1797 Penrith NSW 2751 Australia
4. Hawkesbury Institute for the Environment; Western Sydney University; Locked Bag 1797 Penrith NSW 2751 Australia