1. Department of Watershed Sciences, Utah State University, 5210 Old Main Hill, Logan, UT 84322-5210, USA.
2. Utah Water Research Laboratory, Department of Civil and Environmental Engineering, Utah State University, 8200 Old Main Hill, Logan, UT 84322-8200, USA.
3. US Geological Survey, Utah Cooperative Fish and Wildlife Research Unit, Department of Watershed Sciences, The Ecology Center, Utah State University, 5210 Old Main Hill, Logan, UT 84322-5210, USA.