1. Ecology Department, Fish and Wildlife Ecology and Management Program, Montana State University, Bozeman, MT 59715-5777, USA
2. U.S. Geological Survey, Northern Rocky Mountain Science Center, 2327 University Way, Suite 2, Bozeman, MT 59715, USA
3. Cramer Fish Sciences, Watershed Sciences Lab, 1125 12 Ave. NW Suite B-1, Issaquah, WA 98027, USA