1. Department of Biology, McMaster University, Hamilton, ON, Canada
2. Natural Resources Canada, Canadian Forest Service, Atlantic Forestry Centre, Fredericton, NB, Canada
3. Faculty of Forestry and Environmental Management, University of New Brunswick, Fredericton, NB, Canada
4. Natural Resources Canada, Canadian Forest Service, Great Lakes Forestry Centre, 1219 Queen St. East, Sault Ste. Marie, ON P6A 2E5, Canada