1. British Columbia Ministry of Environment, Conservation Science Section, and the UBC Institute for the Oceans and Fisheries The University of British Columbia, 2202 Main Mall, Vancouver, BC V6T 1Z4, Canada.
2. Pearson Ecological, 2840 Lougheed Highway, Aggasiz, BC VOM 1A1, Canada.
3. Department of Geography, The University of British Columbia, 1984 West Mall, Vancouver, BC V6T 1Z2, Canada.
4. Department of Zoology, The University of British Columbia, 6270 University Boulevard, Vancouver, BC V6T 1Z4, Canada.