1. Institute for the Oceans and Fisheries, Aquatic Ecosystems Research Lab, The University of British Columbia, 2202 Main Mall, Vancouver, BC V6T 1Z4, Canada.
2. Fisheries and Oceans Canada, Gulf Fisheries Centre, Moncton, NB E1C 9B6, Canada.
3. School of Resource and Environmental Management, Simon Fraser University, Burnaby, BC V5A 1S6, Canada.
4. Fisheries and Oceans Canada, Maurice Lamontage Institute, Mont-Joli, QC G5H 3Z4, Canada.