1. Ecology Department, Fish and Wildlife Ecology and Management Program, Montana State University, Bozeman, MT 59717, USA.
2. US Geological Survey, Northern Rocky Mountain Science Center, 2327 University Way, Bozeman, MT 59715, USA.
3. Montana Fish, Wildlife and Parks, 1420 E. Sixth Avenue, Helena, MT 59620, USA.
4. Montana Fish, Wildlife and Parks, 3201 Spurgin Road, Missoula, MT 59804, USA.