1. Evolutionary Biology and Ecology, Free University of Brussels/Université Libre de Bruxelles, avenue FD Roosevelt 50 CP 160/12, B-1050 Brussels, Belgium.
2. Department of Biology, The University of Western Ontario, London ON N6A 5B7, Canada.
3. Institute of Systematic Botany, University of Zürich, Zollikerstrasse 107, CH-8008 Zürich, Switzerland.