1. Department of Biology, Mount Allison University, Sackville, NB E4L 1G7, Canada.
2. Department of Mathematics and Statistics, University of New Brunswick, Fredericton, NB E3B 5A3, Canada.
3. Idaho Department of Fish and Game, 2885 W. Kathleen Avenue, Coeur d’Alene, ID 83835, USA.
4. Golder Associates Ltd., 102, 2535 - 3rd Avenue SE, Calgary, AB T2A 7W5, Canada.